Aizaz, M., Ahmad, W., Asaf, S., Khan, I., Jan, S., Alamri, S., Bilal, S., Jan, R., Kim, K. and Al-Harrasi, A. (2023). Characterization of the seed biopriming, plant growth-promoting and salinity-ameliorating potential of halophilic fungi isolated from hypersaline habitats. Int. J. Mol. Sci. 24: doi:10.3390/ijms24054904.
Amir, M., Prasad, D., Khan, F., Khan, A., Ahamd, B. and Astha. (2024). Seed priming: an overview of techniques, mechanisms, and applications. Plant Sci. Today 11: 553-63. doi:10.14719/pst.2828.
Choi, J., Ju, Y., Nakamichi, A., Cho, S., Woo, S. and Sakagami, J. (2024). Effect of seed hydropriming on the elongation of plumule and radicle during the germination process and changes in enzyme activity under water-deficient conditions. Plants 13: doi:10.3390/plants13243537.
Dela Cruz, S., Rosales, R., Balbas, C., Pascua, G. and Gregorio-Balbas, M. (2026). Hormonal and hydropriming methods enhanced the seedling growth of squash. J. Glob. Innov. Agric. Sci. 14. doi:10.22194/JGIAS/26.1861.
Dela Cruz, S., Rosales, R., Gregorio-Balbas, M., Balbas, C. and Pascua, G. (2025). Effect of seed priming in squash on germination and seedling vigor. Crop Res. 60: 1791-84. doi:10.31830/2454-1761.2025.CR-1015.
Feng, Y., Xiong, J., Lin, G., Cheng, L., Zhang, T., Lin, B., Ke, Z. and Li, X. (2024). Exploring IAA biosynthesis and plant growth promotion mechanism for tomato root endophytes with incomplete IAA synthesis pathways. Chem. Biol. Technol. Agric. 11: doi:10.1186/s40538-024-00712-8.
Fiodor, A., Ajijah, N., Dziewit, L. and Pranaw, K. (2023). Biopriming of seed with plant growth-promoting bacteria for improved germination and seedling growth. Front. Microbiol. 14. doi:10.3389/fmicb.2023.1142966.
Forti, C., Ottobrino, V., Doria, E., Bassolino, L., Toppino, L., Rotino, G., Pagano, A., Macovei, A. and Balestrazzi, A. (2021). Hydropriming applied on fast germinating Solanum villosum Miller seeds: impact on pre-germinative metabolism. Front. Plant Sci. 12: doi:10.3389/fpls.2021. 639336.
Forti, C., Ottobrino, V., Bassolino, L., Toppino, L., Rotino, G., Pagano, A., Macovei, A. and Balestrazzi, A. (2020). Molecular dynamics of pre-germinative metabolism in primed eggplant (Solanum melongena L.) seeds. Hortic. Res. 7. doi:10.1038/s41438-020-0310-8.
ISTA (2008). International rules for seed testing edition. Bassersdorf. The International Seed Testing Association, Richtiarkade 18, CH-8304 Wallisellen, Switzerland.
Kou, X., Han, W. and Kang, J. (2022). Response of root system architecture to water stress at multiple levels: a meta-analysis of trials under controlled conditions. Front. Plant Sci. 13: 1085409. doi:10.3389/fpls.2022.1085409.
Ledina, T., Dordevic, J., Glisic, M., Cobanovic, N., Kovandzic, M. and Bulajic, S. (2024). Technological and functional characteristics of lactic acid bacteria from traditional Serbian chesses. Foods 14: doi:10.3390/foods14010038.
Liu, M., Zhang, Q., Jin, R., Zhao, P., Zhu, X., Wang, J., Yu, Y. and Tang, Z. (2023). The role of IAA in regulating root architecture of sweetpotato (Ipomoea batatas [L.] Lam) in response to potassium deficiency stress. Plants 12: doi:10.3390/plants12091779.
Macdonald, M. and Mohan, V. 2025. Chemical seed priming: molecules and mechanisms for enhancing plant germination, growths and stress tolerance. Curr. Issues Mol. Biol. 47: doi:10.3390/cimb47030177.
Miljakovic, D., Marinkovic, J., Tamindzic, G., Dordevic, V., Tintor, B., Milosevic, D., Ignjatov, M. and Nikolic, Z. (2022). Bio-priming of soybean with Bradyrhizobium japonicum and Bacillus megaterium: strategy to improve seed germination and the initial seedling growth. Plants 11: doi:10.3390/plants11151927.
Nie, L., Song, S., Yin, Q., Zhao, T., Liu, H., He, A. and Wang, W. (2022). Enhancement in seed priming-induced starch degradation of rice seed under chilling stress via GA-mediated α-amylase expression. Rice 15. doi:10.1186/s12284-022-00567-3.
Singh, M., Dotaniya, M., Mishra, A., Dotaniya, C., Regar, K. and Lata, M. (2016). Role of biofertilizers in conservation agriculture. In: Springer Ebooks. Conservation Agriculture. Eds. Busht, J., Meena, V., Mishra, P. and Pattanayak, A. pp.113-34. doi:10.1007/978-981-10-2558-7_4.
Tacbian, D., Rosales, R., Balbas, C., Pascua, g., Bangayan, J., Gregorio-Balbas, M. and Pungtilan, A. (2026). Influence of seed priming agents on germination and seedling vigour in cucumber (Cucumis sativus). Crop Res. 61:165-72. doi:10.31830/2454-1761.2026.CR-1094.
Tamindzic, G., Miljakovic, D., Ignjatov, M., Miladinovic, J., Dordevic, V., Milosevic, D., Jovicic, D., Vlajic, S., Budakov, D. and Grahovac, M. (2024). Impact of simultaneous nutrient priming and biopriming on soybean seed quality and health. Plants 13: doi:10.3390/plants13182557.
Wang, X., Li, J., Ban, L., Wu, Y., Wu, X., Wang, Y., Wen, H., Chapurin, V., Dzyubenko, N., Li, Z., Wang, Z. and Gao, H. (2017). Functional characterization of a gibberellin receptor and its application in alfalfa biomass improvement. Sci. Rep. 7: doi:10.1038/srep41296.
Xu, H., Mo, D., Zhang, X., Li, F., Tao, J., Ge, P., Yang, Y., Wang, Z. and Zhang, Y. (2024). Gibberellin treatment accelerates starch decomposition and seed germination in sticky nightshade (Solanum sisymbriifolium Lam.). Hortic. 10: doi:10.3390/horticulturae10121342.
Yildirim, K., Orel, D., Okyay, H., Gursan, M. and Demir, I. (2021). Quality of immature and mature pepper (Capsicum annuum L.) seeds in relation to bio-priming with endophytic Pseudomonas and Bacillus spp. Hortic. 7: doi:10.3390/horticulturae7040075.
Zhao, T., Deng, X., Xiao, Q., Han, Y., Zhu, S. and Chen, J. (2020). IAA priming improves the germination and seedling growth in cotton (Gossypium hirsutum L.) via regulating the endogenous phytohormones and enhancing the sucrose metabolism. Ind. Crop Prod. 155: doi:10.1016/j. indcrop.2020.112788.
Amir, M., Prasad, D., Khan, F., Khan, A., Ahamd, B. and Astha. (2024). Seed priming: an overview of techniques, mechanisms, and applications. Plant Sci. Today 11: 553-63. doi:10.14719/pst.2828.
Choi, J., Ju, Y., Nakamichi, A., Cho, S., Woo, S. and Sakagami, J. (2024). Effect of seed hydropriming on the elongation of plumule and radicle during the germination process and changes in enzyme activity under water-deficient conditions. Plants 13: doi:10.3390/plants13243537.
Dela Cruz, S., Rosales, R., Balbas, C., Pascua, G. and Gregorio-Balbas, M. (2026). Hormonal and hydropriming methods enhanced the seedling growth of squash. J. Glob. Innov. Agric. Sci. 14. doi:10.22194/JGIAS/26.1861.
Dela Cruz, S., Rosales, R., Gregorio-Balbas, M., Balbas, C. and Pascua, G. (2025). Effect of seed priming in squash on germination and seedling vigor. Crop Res. 60: 1791-84. doi:10.31830/2454-1761.2025.CR-1015.
Feng, Y., Xiong, J., Lin, G., Cheng, L., Zhang, T., Lin, B., Ke, Z. and Li, X. (2024). Exploring IAA biosynthesis and plant growth promotion mechanism for tomato root endophytes with incomplete IAA synthesis pathways. Chem. Biol. Technol. Agric. 11: doi:10.1186/s40538-024-00712-8.
Fiodor, A., Ajijah, N., Dziewit, L. and Pranaw, K. (2023). Biopriming of seed with plant growth-promoting bacteria for improved germination and seedling growth. Front. Microbiol. 14. doi:10.3389/fmicb.2023.1142966.
Forti, C., Ottobrino, V., Doria, E., Bassolino, L., Toppino, L., Rotino, G., Pagano, A., Macovei, A. and Balestrazzi, A. (2021). Hydropriming applied on fast germinating Solanum villosum Miller seeds: impact on pre-germinative metabolism. Front. Plant Sci. 12: doi:10.3389/fpls.2021. 639336.
Forti, C., Ottobrino, V., Bassolino, L., Toppino, L., Rotino, G., Pagano, A., Macovei, A. and Balestrazzi, A. (2020). Molecular dynamics of pre-germinative metabolism in primed eggplant (Solanum melongena L.) seeds. Hortic. Res. 7. doi:10.1038/s41438-020-0310-8.
ISTA (2008). International rules for seed testing edition. Bassersdorf. The International Seed Testing Association, Richtiarkade 18, CH-8304 Wallisellen, Switzerland.
Kou, X., Han, W. and Kang, J. (2022). Response of root system architecture to water stress at multiple levels: a meta-analysis of trials under controlled conditions. Front. Plant Sci. 13: 1085409. doi:10.3389/fpls.2022.1085409.
Ledina, T., Dordevic, J., Glisic, M., Cobanovic, N., Kovandzic, M. and Bulajic, S. (2024). Technological and functional characteristics of lactic acid bacteria from traditional Serbian chesses. Foods 14: doi:10.3390/foods14010038.
Liu, M., Zhang, Q., Jin, R., Zhao, P., Zhu, X., Wang, J., Yu, Y. and Tang, Z. (2023). The role of IAA in regulating root architecture of sweetpotato (Ipomoea batatas [L.] Lam) in response to potassium deficiency stress. Plants 12: doi:10.3390/plants12091779.
Macdonald, M. and Mohan, V. 2025. Chemical seed priming: molecules and mechanisms for enhancing plant germination, growths and stress tolerance. Curr. Issues Mol. Biol. 47: doi:10.3390/cimb47030177.
Miljakovic, D., Marinkovic, J., Tamindzic, G., Dordevic, V., Tintor, B., Milosevic, D., Ignjatov, M. and Nikolic, Z. (2022). Bio-priming of soybean with Bradyrhizobium japonicum and Bacillus megaterium: strategy to improve seed germination and the initial seedling growth. Plants 11: doi:10.3390/plants11151927.
Nie, L., Song, S., Yin, Q., Zhao, T., Liu, H., He, A. and Wang, W. (2022). Enhancement in seed priming-induced starch degradation of rice seed under chilling stress via GA-mediated α-amylase expression. Rice 15. doi:10.1186/s12284-022-00567-3.
Singh, M., Dotaniya, M., Mishra, A., Dotaniya, C., Regar, K. and Lata, M. (2016). Role of biofertilizers in conservation agriculture. In: Springer Ebooks. Conservation Agriculture. Eds. Busht, J., Meena, V., Mishra, P. and Pattanayak, A. pp.113-34. doi:10.1007/978-981-10-2558-7_4.
Tacbian, D., Rosales, R., Balbas, C., Pascua, g., Bangayan, J., Gregorio-Balbas, M. and Pungtilan, A. (2026). Influence of seed priming agents on germination and seedling vigour in cucumber (Cucumis sativus). Crop Res. 61:165-72. doi:10.31830/2454-1761.2026.CR-1094.
Tamindzic, G., Miljakovic, D., Ignjatov, M., Miladinovic, J., Dordevic, V., Milosevic, D., Jovicic, D., Vlajic, S., Budakov, D. and Grahovac, M. (2024). Impact of simultaneous nutrient priming and biopriming on soybean seed quality and health. Plants 13: doi:10.3390/plants13182557.
Wang, X., Li, J., Ban, L., Wu, Y., Wu, X., Wang, Y., Wen, H., Chapurin, V., Dzyubenko, N., Li, Z., Wang, Z. and Gao, H. (2017). Functional characterization of a gibberellin receptor and its application in alfalfa biomass improvement. Sci. Rep. 7: doi:10.1038/srep41296.
Xu, H., Mo, D., Zhang, X., Li, F., Tao, J., Ge, P., Yang, Y., Wang, Z. and Zhang, Y. (2024). Gibberellin treatment accelerates starch decomposition and seed germination in sticky nightshade (Solanum sisymbriifolium Lam.). Hortic. 10: doi:10.3390/horticulturae10121342.
Yildirim, K., Orel, D., Okyay, H., Gursan, M. and Demir, I. (2021). Quality of immature and mature pepper (Capsicum annuum L.) seeds in relation to bio-priming with endophytic Pseudomonas and Bacillus spp. Hortic. 7: doi:10.3390/horticulturae7040075.
Zhao, T., Deng, X., Xiao, Q., Han, Y., Zhu, S. and Chen, J. (2020). IAA priming improves the germination and seedling growth in cotton (Gossypium hirsutum L.) via regulating the endogenous phytohormones and enhancing the sucrose metabolism. Ind. Crop Prod. 155: doi:10.1016/j. indcrop.2020.112788.










