Al-Mayahi, A. M. W. (2021). The effect of humic acid (HA) and zinc oxide nanoparticles (ZnO-NPS) on in vitro regeneration of date palm (Phoenix dactylifera L.) cv. Quntar. Plant Cell Tissue Organ Cult. 145: 445-56. doi:10.1007/s11240-021-02068-5
Akter, S., Sarkar, B. K., Hasan, M. M., Ahmmed, F., Nasrin, P., Al Shajib, G. M., Uddin, S. J., Zaman, F., Kundu, S. K. and Shilpi, J. A. (2025). Evaluation of antioxidant and liver protective activity of ethanolic extract of Vitex peduncularis wall (Lamiaceae) stem bark against carbon tetrachloride (CCl₄)-induced liver damage using in vitro, in vivo, and in silico approaches. Phytomedicine Plus. 5: doi:10.1016/j.phyplu. 2025.100867.
Al-Huqail, A. A., Ali, H. M., Khan, M. N., and Siddiqui, M. H. (2022). The application of silver nanoparticles (AgNPs) improves ex vitro acclimatization and survival rate of micropropagated strawberry (Fragaria × ananassa Duch.) plantlets. Horticulturae 8: doi.org/10.3390/horticulturae8060524.
Bakhtiar, Z., Khalili, F. A., Ghasemi, M. and Mirjalili, M. H. (2025). Micropropagation, callus induction, and cell culture establishment of Zataria multiflora (Lamiaceae): An efficient biotechnological platform for the production of Rosmarinic acid. Ind. Crops Prod. 226: doi:10.1016/j.indcrop.2025.120701.
Blanco-Murillo, F., Marín-Guirao, L., Rodríguez-Rojas, F., Sola, I., Carbonell-Garzón, E., Sánchez-Lizaso, J. L. and Sáez, C. A. (2024). The evolutive role of shoot apical meristems in the adaptation of angiosperms to life at sea and the jump to potential environmental biotechnology applications. Sci. Total Environ. 2024: doi:10.1016/j. scitotenv.2024.176917.
Chaouch, R., Soufi, S., Kthiri, Z. and Bettaieb, T. (2025). Auxin-like potential of essential oils in strawberry micropropagation: A sustainable approach to root induction and photosynthetic performance. South Afr. J. Bot. 177: 352-62. doi:10.1016/j. sajb.2024.12.025.
Csorba, C., Rodić, N., Zhao, Y., Antonielli, L., Brader, G., Vlachou, A., Tsiokanos, E., Lalaymia, I., Declerck, S., Papageorgiou, V. P., Assimopoulou, A. N. and Sessitsch, A. (2022). Metabolite production in alkanna tinctoria links plant development with the recruitment of individual members of microbiome thriving at the root-soil interface. mSystems 7: doi:10.1128/msystems.00451-22.
Direk, A., Arikan-Abdulveli, B., Balci, M., Gurkan, E., Gulenturk, E. O., Ozfidan-Konakci, C. and Yildiztugay, E. (2026). Biochar as an alternative strategy to mitigate europium oxide nanoparticle toxicity in maize: Improvements in water status, ion homeostasis, photosynthetic electron flux, and antioxidant defense. Environ. Res. 299: doi.org/10.1016/j.envres.2026.124363.
Ghorbanpour, M., Movahedi, A., Hatami, M., Kariman, K., Bovand, F., and Shahid, M. A. (2021). Insights into nanoparticle-induced changes in plant photosynthesis. Photosynthetica 59: 570-86. doi.org/10.32615/ps.2021.049.
Hong, X., Zhang, H. and Yang, S. (2026). Natural products and biological activity from fungal symbionts associated with Rosmarinus officinalis L. Nat. Prod. Res. 38: 1-5. doi:10.1080/14786419.2026.2638953.
John, S., Apelt, F., Kumar, A., Acosta, I. F., Bents, D., Annunziata, M. G., Fichtner, F., Gutjahr, C., Mueller-Roeber, B. and Olas, J. J. (2024). The transcription factor HSFA7b controls thermomemory at the shoot apical meristem by regulating ethylene biosynthesis and signaling in Arabidopsis. Plant Comm. 5: doi:10.1016/j.xplc.2023. 100743.
Lala, S. (2021). Nanoparticles as elicitors and harvesters of economically important secondary metabolites in higher plants: A review. IET Nanobiotechnology 15: 28-57. doi:10.1049/nbt2.12005.
Mamdouh, D., and Smetanska, I. (2022). Optimization of callus and cell suspension cultures of Lycium schweinfurthii for improved production of phenolics, flavonoids, and antioxidant activity. Horticulturae 8: doi.org/10.3390/horticulturae8050394.
Mawed, S. A., Ghoneim, M. H., Farag, M. R., Abdelmoety, M. S. H., Ismail, T. A. and Al-Zahaby, S. A. (2024). Aluminium nanoparticles (AL-NPs) caused gastrointestinal malformations, metabolic disorders and steatohepatitis in zebrafish larvae: Modulatory role of Rosmarinic acid. Aquaculture Rep. 39: doi:10.1016/j.aqrep.2024.102489.
Nartop, P., and Özdemir, F. A. (2022). The effects of green-synthesized silver nanoparticles on the ex vitro acclimatization and survival rate of plantlets. Turk. J. Agric. For. 46: 485-96. doi.org/10.3906/tar-2111-53.
Ozyigit, I. I., Dogan, I., Hocaoglu-Ozyigit, A., Yalcin, B., Erdogan, A., Yalcin, I. E., Cabi, E. and Kaya, Y. (2023). Production of secondary metabolites using tissue culture-based biotechnological applications. Front. Plant Sci. 14: doi:10.3389/fpls.2023. 1132555.
Vashishta, P. and Asija, R. (2026). Development of a Rosmarinus officinalis extract-loaded nanoemulgel for topical anti-inflammatory delivery: Molecular docking, HPTLC profiling, and ex vivo evaluation. Next Nanotechnol. 9: doi:10.1016/j.nxnano.2026. 100418.
Rivera-Méndez, W., Obregón, J., Morán-Morales, J. and de la Cruz Chacón, I. (2023). Nano-elicitors: A new trend in plant secondary metabolism induction. Plants 9: doi:10. 3390/plants9091123.
Saeed, F., Younas, M., Fazal, H., Mushtaq, S., Rahman, F. U., Shah, M., Anjum, S., Ahmad, N., Ali, M., Hano, C., & Abbasi, B. H. (2021). Green and chemically synthesized zinc oxide nanoparticles: Effects on in‑vitro seedlings and callus cultures of Silybum marianum and evaluation of their antimicrobial and anticancer potential. Artif. Cells Nanomed. Biotechnol. 49: 450-60. doi:10.1080/21691401.2021. 1926274.
Silva, G. M., Campos, E. V. R., de Oliveira, F. F., de Souza Rodrigues, J., de Freitas Proença, P. L., Melo, A. A. and Fraceto, L. F. (2025). Harnessing nanotechnology and bio-based agents: Advanced strategies for sustainable soybean nematode management. Plant Nano Biol. 13: doi:10.1016/j.plana.2025.100195.
Singh, S. and Agrawal, N. (2024). Exploring the pharmacological potential and bioactive components of Pogostemon cablin (Blanco) Benth, traditional Chinese medicine. Pharmacol. Res. Mod. Chin. Med. 10: doi:10.1016/j.prmcm.2024.100382.
Singleton, V. L. and Rossi, J. A. (1965). Colorimetry of total phenolics with phosphomolybdic-phosphotungstic acid reagents. Am. J. Enol. Vitic. 16: 144–58. doi:10.5344/ajev.1965.16. 3.144.
Tymoszuk, A. and Kulus, D. (2025). The effect of zinc oxide nanoparticles on the quantitative and qualitative traits of Scutellaria baicalensis Georgi in in vitro culture. Int. J. Mol. Sci. 26: doi:10.3390/ijms26125836.
Zamanian, M., Azizi, M. H. and Thibault, J. (2024). Development of bio-based PVA/PEG nanocomposite films incorporating nanoencapsulated rosemary (Rosmarinus officinalis L.) essential oil for industrial non-food applications. J. Agric. Food Chem. 72: 8805-816. doi:10.1016/j.polymertesting.2026.109206.
Zayed, M. S., Ahmed, R. and Hassan, S. (2024). Nanoparticles enhance in vitro micropropagation and secondary metabolite accumulation in Origanum petraeum. Nanomaterials 15: doi:10.3390/nano15191496.
Akter, S., Sarkar, B. K., Hasan, M. M., Ahmmed, F., Nasrin, P., Al Shajib, G. M., Uddin, S. J., Zaman, F., Kundu, S. K. and Shilpi, J. A. (2025). Evaluation of antioxidant and liver protective activity of ethanolic extract of Vitex peduncularis wall (Lamiaceae) stem bark against carbon tetrachloride (CCl₄)-induced liver damage using in vitro, in vivo, and in silico approaches. Phytomedicine Plus. 5: doi:10.1016/j.phyplu. 2025.100867.
Al-Huqail, A. A., Ali, H. M., Khan, M. N., and Siddiqui, M. H. (2022). The application of silver nanoparticles (AgNPs) improves ex vitro acclimatization and survival rate of micropropagated strawberry (Fragaria × ananassa Duch.) plantlets. Horticulturae 8: doi.org/10.3390/horticulturae8060524.
Bakhtiar, Z., Khalili, F. A., Ghasemi, M. and Mirjalili, M. H. (2025). Micropropagation, callus induction, and cell culture establishment of Zataria multiflora (Lamiaceae): An efficient biotechnological platform for the production of Rosmarinic acid. Ind. Crops Prod. 226: doi:10.1016/j.indcrop.2025.120701.
Blanco-Murillo, F., Marín-Guirao, L., Rodríguez-Rojas, F., Sola, I., Carbonell-Garzón, E., Sánchez-Lizaso, J. L. and Sáez, C. A. (2024). The evolutive role of shoot apical meristems in the adaptation of angiosperms to life at sea and the jump to potential environmental biotechnology applications. Sci. Total Environ. 2024: doi:10.1016/j. scitotenv.2024.176917.
Chaouch, R., Soufi, S., Kthiri, Z. and Bettaieb, T. (2025). Auxin-like potential of essential oils in strawberry micropropagation: A sustainable approach to root induction and photosynthetic performance. South Afr. J. Bot. 177: 352-62. doi:10.1016/j. sajb.2024.12.025.
Csorba, C., Rodić, N., Zhao, Y., Antonielli, L., Brader, G., Vlachou, A., Tsiokanos, E., Lalaymia, I., Declerck, S., Papageorgiou, V. P., Assimopoulou, A. N. and Sessitsch, A. (2022). Metabolite production in alkanna tinctoria links plant development with the recruitment of individual members of microbiome thriving at the root-soil interface. mSystems 7: doi:10.1128/msystems.00451-22.
Direk, A., Arikan-Abdulveli, B., Balci, M., Gurkan, E., Gulenturk, E. O., Ozfidan-Konakci, C. and Yildiztugay, E. (2026). Biochar as an alternative strategy to mitigate europium oxide nanoparticle toxicity in maize: Improvements in water status, ion homeostasis, photosynthetic electron flux, and antioxidant defense. Environ. Res. 299: doi.org/10.1016/j.envres.2026.124363.
Ghorbanpour, M., Movahedi, A., Hatami, M., Kariman, K., Bovand, F., and Shahid, M. A. (2021). Insights into nanoparticle-induced changes in plant photosynthesis. Photosynthetica 59: 570-86. doi.org/10.32615/ps.2021.049.
Hong, X., Zhang, H. and Yang, S. (2026). Natural products and biological activity from fungal symbionts associated with Rosmarinus officinalis L. Nat. Prod. Res. 38: 1-5. doi:10.1080/14786419.2026.2638953.
John, S., Apelt, F., Kumar, A., Acosta, I. F., Bents, D., Annunziata, M. G., Fichtner, F., Gutjahr, C., Mueller-Roeber, B. and Olas, J. J. (2024). The transcription factor HSFA7b controls thermomemory at the shoot apical meristem by regulating ethylene biosynthesis and signaling in Arabidopsis. Plant Comm. 5: doi:10.1016/j.xplc.2023. 100743.
Lala, S. (2021). Nanoparticles as elicitors and harvesters of economically important secondary metabolites in higher plants: A review. IET Nanobiotechnology 15: 28-57. doi:10.1049/nbt2.12005.
Mamdouh, D., and Smetanska, I. (2022). Optimization of callus and cell suspension cultures of Lycium schweinfurthii for improved production of phenolics, flavonoids, and antioxidant activity. Horticulturae 8: doi.org/10.3390/horticulturae8050394.
Mawed, S. A., Ghoneim, M. H., Farag, M. R., Abdelmoety, M. S. H., Ismail, T. A. and Al-Zahaby, S. A. (2024). Aluminium nanoparticles (AL-NPs) caused gastrointestinal malformations, metabolic disorders and steatohepatitis in zebrafish larvae: Modulatory role of Rosmarinic acid. Aquaculture Rep. 39: doi:10.1016/j.aqrep.2024.102489.
Nartop, P., and Özdemir, F. A. (2022). The effects of green-synthesized silver nanoparticles on the ex vitro acclimatization and survival rate of plantlets. Turk. J. Agric. For. 46: 485-96. doi.org/10.3906/tar-2111-53.
Ozyigit, I. I., Dogan, I., Hocaoglu-Ozyigit, A., Yalcin, B., Erdogan, A., Yalcin, I. E., Cabi, E. and Kaya, Y. (2023). Production of secondary metabolites using tissue culture-based biotechnological applications. Front. Plant Sci. 14: doi:10.3389/fpls.2023. 1132555.
Vashishta, P. and Asija, R. (2026). Development of a Rosmarinus officinalis extract-loaded nanoemulgel for topical anti-inflammatory delivery: Molecular docking, HPTLC profiling, and ex vivo evaluation. Next Nanotechnol. 9: doi:10.1016/j.nxnano.2026. 100418.
Rivera-Méndez, W., Obregón, J., Morán-Morales, J. and de la Cruz Chacón, I. (2023). Nano-elicitors: A new trend in plant secondary metabolism induction. Plants 9: doi:10. 3390/plants9091123.
Saeed, F., Younas, M., Fazal, H., Mushtaq, S., Rahman, F. U., Shah, M., Anjum, S., Ahmad, N., Ali, M., Hano, C., & Abbasi, B. H. (2021). Green and chemically synthesized zinc oxide nanoparticles: Effects on in‑vitro seedlings and callus cultures of Silybum marianum and evaluation of their antimicrobial and anticancer potential. Artif. Cells Nanomed. Biotechnol. 49: 450-60. doi:10.1080/21691401.2021. 1926274.
Silva, G. M., Campos, E. V. R., de Oliveira, F. F., de Souza Rodrigues, J., de Freitas Proença, P. L., Melo, A. A. and Fraceto, L. F. (2025). Harnessing nanotechnology and bio-based agents: Advanced strategies for sustainable soybean nematode management. Plant Nano Biol. 13: doi:10.1016/j.plana.2025.100195.
Singh, S. and Agrawal, N. (2024). Exploring the pharmacological potential and bioactive components of Pogostemon cablin (Blanco) Benth, traditional Chinese medicine. Pharmacol. Res. Mod. Chin. Med. 10: doi:10.1016/j.prmcm.2024.100382.
Singleton, V. L. and Rossi, J. A. (1965). Colorimetry of total phenolics with phosphomolybdic-phosphotungstic acid reagents. Am. J. Enol. Vitic. 16: 144–58. doi:10.5344/ajev.1965.16. 3.144.
Tymoszuk, A. and Kulus, D. (2025). The effect of zinc oxide nanoparticles on the quantitative and qualitative traits of Scutellaria baicalensis Georgi in in vitro culture. Int. J. Mol. Sci. 26: doi:10.3390/ijms26125836.
Zamanian, M., Azizi, M. H. and Thibault, J. (2024). Development of bio-based PVA/PEG nanocomposite films incorporating nanoencapsulated rosemary (Rosmarinus officinalis L.) essential oil for industrial non-food applications. J. Agric. Food Chem. 72: 8805-816. doi:10.1016/j.polymertesting.2026.109206.
Zayed, M. S., Ahmed, R. and Hassan, S. (2024). Nanoparticles enhance in vitro micropropagation and secondary metabolite accumulation in Origanum petraeum. Nanomaterials 15: doi:10.3390/nano15191496.
